I'm not sure to which FM Darryl refers, but Gary's approach does not seem to follow Burnham and Anderson (2002: 332) who tentatively recommend the number of animals (not sum(R_i) or a larger number) as the sample size for survival parameters in fixed effects modelling (see also their extensive chain binomial example on p207 et seq.). B&A also say "The issue of sample size can be complex... We do not pursue solutions here. We just raise the issue as a future research area". Has that research been done since? Not a big deal, but religious use of AICc is less attractive if we cannot agree on n, even to within an order of magnitude.
Murray